PhD Student Position in Biochemistry and Molecular Medicine

University of Oulu



Applications are invited for a full-time PhD student position for a maximum of four years, starting at the earliest on 01.01.2020 in a Biocenter Oulu research project headed by Professor Johanna Myllyharju in the Hypoxia & Collagens research unit, Faculty of Biochemistry and Molecular Medicine.

Our research focuses on the key enzyme regulators of the hypoxia response and collagen synthesis and their potential as therapeutic targets. We use mainly biochemical, cell biological and genetic mouse models to understand how these enzymes regulate normal physiological and developmental processes. In addition, we study whether their inhibition has therapeutic value in selected common pathological conditions, with main focus currently on fibrosis, cancer and inflammation. We offer a research environment with strong experience in PhD training, excellent technological facilities and a friendly and supporting community. More information can be found on our web page


To successfully occupy the position of a doctoral student, the candidate must possess a Master’s degree or an equivalent degree (e.g. licentiate) preferably in the field of biochemistry, cell biology, medical science or related fields. Degree must have been completed with good grades by 31.12.2019. Ability to pursue independent research and excellent writing and fluency in English is expected. The doctoral student position is intended for an Early Stage Researcher (ESR) who, according to the ”European Charter for Researchers and the Code of Conduct for the Recruitment of Researchers”, is defined as a researcher in the first four years (full-time equivalent) of his/her research activity, including the period of research training. Applicants will be either starting their doctoral training or will have completed no more than two full years of training (adjusted for possible previous part-time status) at the time the position is filled. The selected candidate will carry out his/her doctoral studies in the Health and Biosciences Doctoral Programme of the University of Oulu Graduate School (UniOGS) and must meet the minimal requirements to receive doctoral study rights as defined in


The salary of the selected doctoral student will be initially set on level 2, and later on levels 2 – 4 of the national salary scale for the teaching and research staff of Finnish universities. In addition, a supplementary remuneration is given for personal achievement and performance (max. 50 %). In practice, the salary is roughly 2200 – 2700 € per month for an early stage researcher, depending on the appointee’s qualifications and progress of the doctoral thesis.

Since this call is for a starting doctoral researcher, the position is filled for a four-year period, beginning on 1 January 2020. A six-month trial period is applied in the position.


Applications must be submitted using the electronic application form by September 16, 2019, 24:00 (Finnish local time), with the following attachments as three separate pdf files:

  1. Complete CV containing contact information of at least two referees (Max. size limit of the attachment: 5 Mbytes)
  2. Motivation letter (maximum 1 page) describing prior knowledge and research interests and career plans (1-2 pages; max. size limit of the attachment: 2 Mbytes).
  3. Certificates/Diplomas: Scanned electronic copies of diplomas and transcripts of the records of relevant previous degrees. If the original documents are not in English, Finnish or Swedish, each document must be accompanied by an official certified translation into English or Finnish. (Max. size limit of the attachment: 2 Mbytes)

Applications sent by email will not be considered.

Other Relevant Information

The eligible applicants fitting best in the research profile expected for the position may be invited to an on-site or remote interview. All applicants will be notified when the selection process is completed.


Prof. Johanna Myllyharju, e-mail: johanna.myllyharju (at)

In your application, please refer to


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